The Echinoid Directory

• Apical disc relatively large with pentastellate pattern of narrow ridges best developed in juveniles (ridges can become more or less obliterated in very large individuals).
• Periproct large and diamond-shaped, larger than suranal plate.
• Ambulacral zones very narrow; bigeminate adorally and either bigeminate or unigeminate at ambitus and aborally.
• Deep circular sphaeridial pits clearly developed close to every second pore-pair adorally.
• Primary interambulacral tubercles large and circular; mamelon relatively small adorally and increasing in size adapically.

Upper Albian to Cenomanian of the U.S.A., Europe and the Middle East.

Goniophorus lunulatus L. Agassiz, 1838, by subsequent designation of Lambert & Thiery, 1911, p. 209.

• Goniophorus scotti Lambert, 1927; Upper Albian to Cenomanian of Texas, U.S.A. and Upper Albian of Kent, England [includes Goniophorus whitneyi Ikins (1940)].
• Goniophorus lunulatus L. Agassiz, 1838: Late Albian S. dispar Zone to Early Cenomanian, M. dixoni Zone, Southern England and Northern France.
• Goniophorus lamberti (de Loriol, 1909): Cenomanian of Syria.

Salenioida; Saleniidae; Hyposaleniinae.

Monophyletic.

Distinguished from Hyposalenia by its characteristic disc ornamentation and the presence of obvious sphaeridial pits in ambulacral zones.

• Goniophorus? pentagonalis Muller, 1859, p. 6, pl. 7, fig. 3 is probably not a calycinid. See Ham (1987).
• Goniophorus lorioli Lambert & Thiery 1911, p. 209, from the Albian of Basses-Alpes, France is indeterminate.