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Upper Cretaceous (Campanian - Maastrichtian) North Africa, Iran, Arabian Peninsula.
Remarks
Gauthier, V. 1889. Exploration scientifique de la Tunisie. Illustrations de la partie Paleontologique et geologique. Fascicule 1. Echinides Fossiles. Paris: Imprimerie nationale.
Smith, A. B. 1995. Late Campanian - Maastrichtian echinoids from the United Arab Emirates - Oman borders region. Bulletin of the Natural History Museum London (Geology Series)51, 121-240.