Brachycera (Cyclorrhapha – Calyptrata)

The Calyptrata group contain some of the most well known flies eg house flies and blow-flies.

The Calyptrata are distinguished from other Cyclorrhapha by:

  • well-developed calypters, which are membranous flaps situated under the base of the wings
  • a complete dorsal transverse suture on the thorax

Calyptrata are generally rather robust and bristly flies.


Adult bot-flies have reduced mouthparts while tsetse flies have piercing mouthparts for blood-sucking.

Flat-flies, keds and bat-flies are blood-sucking ectoparasites of:

  • bats
  • birds
  • other mammals

Adults of some species are wingless or have reduced wings, others may initially be fully winged but the wings fall off once they have located a suitable host.


The adult fly emerges from the pupal case (puparium) through a circular aperture.

A small membranous flap at the base of the hind edge of the wing in some flies.