Dr Ralf Britz

Image of Dr Ralf Britz
  • Researcher
  • Life Sciences department
  • Vertebrates Division
Natural History Museum
Cromwell Road
London
SW7 5BD

Biography


Academic positions

since 2004       Researcher Fishes, The Natural History Museum, London, UK

05 2012           Visiting professor, Departamento de Zoologia,Universidade de São Paulo, Brazil

05 2010           Visiting professor, Departamento de Zoologia,Universidade de São Paulo, Brazil

07/08 2007      Visiting professor, Departamento de Zoologia, Universidade de São Paulo, Brazil

since 2005       Research Associate, Division of Fishes, NMNH, Smithsonian Institution,
                                                                                                           Washington D.C., USA

09/10 2003      Invited Researcher, Département Systématique et Evolution, MNHN, Paris,
                                                                                                           France

2002-2004       Visiting Researcher, Division of Fishes, NMNH,Smithsonian Institution,
                                                                                                          Washington D.C., USA

1998-2004       Research Associate, Ichthyological Department, AMNH, New York, USA

1996-2004       Assistant Professor, Lehrstuhl für Spezielle Zoologie, Tübingen University,
                                                                                                          Germany
1995-1996       Postdoctoral Fellow, Ichthyological Department, AMNH, New York, USA

1988-1995       Teaching Assistant, Lehrstuhl für Spezielle Zoologie, Tübingen University,
                                                                                                           Germany

 


Education

1992-1995         PhD, Lehrstuhl für Spezielle Zoologie, Tübingen University, Germany
1985-1992         Biologiediplom (Biology Masters), Lehrstuhl für Spezielle Zoologie,
                                                                                                            Tübingen University, Germany


Research


My research program focuses on comparative anatomy, phylogenetics, and taxonomy of actinopterygian and sarcopterygian fishes, ranging from general questions of evolutionary biology and homology and studies of higher level relationships to basal problems in taxonomy and systematics of various fish groups. Although my primary approach to phylogenetic questions is and will remain deeply founded in morphology, I have also developed an interest in tests of morphology-based hypotheses with molecular data and I am involved in several ongoing collaborations.

Weberian apparatus

Developing Weberian apparatus of a 8.4 mm Rhabdalestes larva (image: R. Britz).


My taxonomic interest lies with the freshwater fishes of Southeast Asia with a focus on Myanmar (Burma), a poorly known fauna with dozens of new species described each year. These systematic studies form the basis for more general questions about biogeographical pattern of this interesting region and how these have evolved.

Myanmar fishes

New freshwater fish species discovered in Myanmar and described in the last 10 years by myself and coauthors (images: R. Britz).


 

The morphology based questions I try to resolve concern the relationship between evolution and development. I investigate how ontogenies change during evolution and try to understand how complex structures, like e.g. the tail of the ocean sunfishes or the Weberian apparatus of otophysans, have evolved. In addition, I study the ontogeny of characters to resolve long-standing homology issues and thus to get a better idea about the phylogenetic relationships of various groups.

Paedocypris

Paedocypris progenetica, the smallest fish and vertebrate species, in comparison to a 5 p coin. Cleared and double stained skeleton of male and female Paedocypris (images: R. Britz).


I have recently become fascinated by the organismal consequences of miniaturization in teleosts caused by the evolutionary process of developmental truncation (progenesis) for which Paedocypris spp. and Danionella dracula are prime examples.

Dracula

Danionella dracula, alive (above), scanning electron micrograph of its head (middle) and head of cleared and double stained male (below) illustrating fangs (images: R. Britz).


Publications

112. Britz, R., Conway, K. & L. Rüber. (2014). Miniatures, morphology and molecules: Paedocypris and its phylogenetic position (Teleostei, Cypriniformes). Zoological Journal of the Linnean Society: 60pp. Early View, doi: 10.1111/zoj.12184.

111. Mattox, G., Britz, R. & Toledo-Piza, M. (2014). Skeletal development and ossification sequence of the characiform Salminus brasiliensis (Ostariophysi: Characidae). Ichthyological Exploration of Freshwaters, 25: 103-158.

110. Low, B. W., Tan, H. H. & Britz, R. (2014). Trichopodus poptae, a new anabantoid fish from Borneo (Teleostei: Osphronemidae).  Ichthyological Exploration of Freshwaters, 25: 69-77.

109. Conway, K. W., Britz, R. & Siegel, D. (2014). Different on the inside: extreme swimbladder sexual dimorphism in the South Asian torrent minnows. Biology Letters, 10: http://dx.doi.org/10.1098/rsbl.2014.0348

108. Toledo-Piza, M., Mattox, G. & R. Britz. (2014). Priocharax nanus, a new miniature characid from the rio Negro, Amazon basin (Ostariophysi: Characiformes), with an updated list of miniature Neotropical freshwater fishes. Neotropical Ichthyology, 12: 229-246.

107. Rocha, L., et al. (2014). Specimen collection: an essential tool. Science, 344: 814-815.

106. Britz, R., Rüber, L. & G. D. Johnson. (2014). Reinventing the disc: a reminder to give credit to past giants. Proceedings of the Royal Society B, 281: http://dx.doi.org/10.1098/rspb.2013.2920

105. Britz, R., Kakkassery, F. & R. Raghavan. (2014). Osteology of Kryptoglanis shajii, a stygobitic catfish (Teleostei: Siluriformes) from Peninsular India with a diagnosis of the new family Kryptoglanidae. Ichthyological Exploration of Freshwaters, 24: 193-207.

104. Britz, R. & S. O. Kullander. (2013). Dario kajal, a new species of badid fish from Meghalaya, India (Teleostei: Badidae). Zootaxa, 3731: 331-337.

103. Britz, R. (2013). Channa andrao, a new species of dwarf snakehead from West Bengal, India (Teleostei: Channidae). Zootaxa 3731: 287-294.

102. Britz, R., Mattox, G. M. T., Toledo-Piza, M. & M. M. F. Marinho. (2013). Nomenclatural remarks on Cyanogaster Oken 1823 and Cyanogaster Blainville 1825 (Mollusca: Gastropoda: Pleurobranchaeidae). Zootaxa 3683: 99-100.

101. Mattox, G. M. T., Britz, R., Toledo-Piza, M. & M. M. F. Marinho. (2013). Cyanogaster noctivaga, a remarkable new genus and species of miniature fish from the Rio Negro, Amazon basin (Ostariophysi: Characidae). Ichthyological Exploration of Freshwaters, 23: 297-318.

100. Britz, R., Ali, A. & R. Raghavan. (2013). Pseudolaguvia lapillicola, a new species of catfish from Peninsular India (Teleostei: Sisoridae). Ichthyological Exploration of Freshwaters, 23: 289-295.

99. Britz, R., Ali, A., Siby, P., Kumar, K. & R. Raghavan (2012). First record from the wild of Carinotetraodon imitator in Peninsular India (Teleostei: Tetraodontiformes: Tetraodontidae). Ichthyological Exploration of Freshwaters, 23: 105-109.

98. Britz, R. & K. Hartel. (2012). On the synonymy of Caristius groenlandicus Jensen and Pteraclis fasciatus Borodin (Pisces: Caristiidae). Zootaxa, 3546: 85-88.

97. Britz, R. & M. Toledo-Piza. (2012). Egg surface structure of the freshwater toadfish Thalassophryne amazonica (Teleostei: Batrachoididae) with information on its distribution and natural habitat. Neotropical Ichthyology, 10: 593-599.

96. Britz, R. & G. D. Johnson. (2012). Ontogeny and homology of the skeletal elements that form the sucking disc of remoras (Teleostei, Echeneoidei, Echeneidae). Journal of Morphology, 273: 1353-1356. DOI: 10.1002/jmor.20063

95. Britz, R., Ali, A. & R. Raghavan. (2012). Pangio ammophila, a new species of eel-loach from Karnataka, southern India (Teleostei: Cypriniformes: Cobitidae). Ichthyological Exploration of Freshwaters, 23: 45-50.

94. Britz, R., Ali, A. & S. Philip. (2012). Dario urops, a new species of badid fish from the Western Ghats, southern India (Teleostei: Percomorpha: Badidae). Zootaxa, 3348: 63-68.

93. Britz, R., Kumar, K. & F. Baby. (2012). Pristolepis rubripinnis, a new species of fish from southern India (Teleostei: Percomorpha: Pristolepididae). Zootaxa, 3345: 59-68.

92. Fraser, G. J., Britz, R., Hall, A., Johanson, Z. & M. M. Smith. (2012). Replacing the first-generation dentition in pufferfish with a unique beak. Proceedings of the National Academy of Sciences, 109: 8179-8184. Published on-line 07 May 2012, doi: 10.1073/pnas.1119635109

91. Britz, R. & G. D. Johnson. (2012). The caudal skeleton of a 20 mm Triodon and homology of its components. Proceedings of the Biological Society of Washington, 125: 66-73.

90.  Britz, R., Kottelat, M. & H. H. Tan. (2012). Fangfangia spinicleithralis, a new genus and species of miniature cyprinid from the peat swamp forests of Borneo (Teleostei: Cypriniformes: Cyprinidae). Ichthyological Exploration of Freshwaters, 22: 327-335.

89. Bohlen, J., Slechtova, V., Tan H. H., & R. Britz. (2011). Phylogeny of the Southeast Asian freshwater fish genus Pangio (Cypriniformes; Cobitidae). Molecular Phylogenetics and Evolution, 61: 854-865.

88. Britz, R. & K. W. Conway (2011). Additions to “The Cypriniformes tree of confusion”. Zootaxa, 2946: 142.

87. Britz, R. & K. W. Conway (2011). The Cypriniformes tree of confusion. Zootaxa, 2946: 73-78.

86. Britz, R. & G. D. Johnson (2011). Comments on the establishment of the one to one relationship between characters as a prerequisite for homology assessment in phylogenetic studies. Zootaxa, 2946: 65-72.

85. Britz, R., Lalremsanga, H. T., Lalrotluanga & Lalramliana. (2011). Monopterus ichthyophoides, a new species of scaled swamp eel (Teleostei: Synbranchiformes: Synbranchidae) from Mizoram, India. Zootaxa, 2936: 51-58.

84. Benziger, A.,  Philip, S., Raghavan, R., Ali, P. H. A.,Sukumaran, M., Tharian, J. C., Dahanukar, N., Baby, F., Peter, R., Rema Devi, K., Radhakrishnan, K. V., Haniffa, M. A. K., Britz, R. & Antunes, A. (2011). Unraveling a 146 years old taxonomic puzzle: validation of Malabar Snakehead, species-status and its relevance for channid systematics and evolution. PLOS one, 6(6): e21272. Published online on 24 June 2011, doi:10.1371/journal.pone.0021272

83. Brown, K. J.,  Britz, R., Bills R., Rüber, L. & Day J. J. (2011). Pectoral fin loss in the Mastacembelidae: a new species from Lake Tanganyika. Journal of Zoology, Journal of Zoology, 284: 286-293. Published online on 1 April 2011, doi:10.1111/j.1469-7998.2011.00804.x

82. Vishwanath, W., Ng, H.H., Britz, R., Singh, L.K., Chaudhry, S. & Conway, K. W. (2011). Chapter 3. The status and distribution of freshwater fishes of the Eastern Himalaya region, pp. 22-41, in Allen, D., Molur, S. & Daniel, B. A. (compilers) The Status and distribution of freshwater biodiversity in the eastern Himalaya. Cambridge, UK and Gland, Switzerland: IUCN, & Coimbatore, India: Zoo Outreach Organisation.

81. Britz, R. & Win, T. (2010). Redescription of Silurus burmanensis Thant 1967: a Silurus not a Pterocryptis (Teleostei: Siluriformes: Siluridae). Zootaxa, 2647: 61-68.

80. Conway, K. W. & Britz, R. (2010). Three new species of Psilorhynchus from the Ayeyarwaddy River drainage, Myanmar (Teleostei: Psilorhynchidae). Zootaxa, 2616: 31-47.

79. Britz, R. (2010). A new earthworm eel of the genus Chaudhuria from the Ayeyarwaddy River Drainage, Myanmar (Teleostei: Synbranchiformes: Chaudhuriidae). Zootaxa, 2571: 62-68.

78. Britz, R. (2010). Macrognathus aureus, a new spiny eel of the M. aculeatus species group from the Upper Ayeyarwaddy River Drainage, Myanmar (Teleostei: Synbranchiformes: Mastacembelidae). Zootaxa, 2514: 55-60.

77. Schnell, N., Britz, R. & Johnson, G. D. (2010). New insights into the complex structure and ontogeny of the occipito-vertebral gap in barbeled dragonfishes (Stomiidae, Teleostei). Journal of Morphology, 271:1006–1022. Published on-line 17 May 2010 (DOI: 10.1002/jmor.10858)

76. Britz, R. (2010). Review of MESOZOIC FISHES 4. Edited by Arratia, G., Schultze, H.-P., Wilson, M. V. H. Journal of Fish Biology, 76: 1537-1539.

75. Britz, R. & Kottelat, M. (2010). Pangio longimanus, a miniature species of eel-loach from Central Laos (Teleostei: Cypriniformes: Cobitidae). Ichthyological Exploration of  Freshwaters 20: 371-376

74. Britz, R. (2010). Species of the Macrognathus aculeatus group in Myanmar with remarks on M. caudiocellatus (Teleostei: Synbranchiformes: Mastacembelidae). Ichthyological Exploration of  Freshwaters 20: 295-308.

73. Hilton, E. & Britz, R. (2010). The caudal skeleton of osteoglossomorph fishes, revisited: comparisons, homologies, and characters in Nelson, J. S., Schultze, H.-P. & Wilson, M.V.H. (eds.): Origin and phylogenetic interrelationships of teleosts honoring Gloria Arratia. Verlag Dr Friedrich Pfeil, pp. 219-237.

72. Britz, R. & Johnson, G. D. (2010). Occipito-vertebral fusion in actinopterygians: conjecture, myth and reality. Part 1: Non-teleosts in Nelson, J. S., Schultze, H.-P. & Wilson, M.V.H. (eds.): Origin and phylogenetic interrelationships of teleosts honoring Gloria Arratia. Verlag Dr Friedrich Pfeil, pp. 77-93.

71. Johnson, G. D. & Britz, R. (2010). Occipito-vertebral fusion in actinopterygians: conjecture, myth and reality. Part 2: Teleosts in Nelson, J. S., Schultze, H.-P. & Wilson, M.V.H. (eds.): Origin and phylogenetic interrelationships of teleosts honoring Gloria Arratia. Verlag Dr Friedrich Pfeil, pp. 95-110.

70. Britz, R. (2009). Teleostei in Westheide, W. & Rieger, R.(eds.): Spezielle Zoologie, Teil 2: Wirbeltiere. 2te Auflage. Gustav Fischer Verlag, pp. 260-306. (Chapter Teleostei in Volume 2 [Vertebrate systematics], second edition of the major German zoology textbook)

69. Britz, R. (2009). Danionella priapus, a new species of miniature cyprinid fish from West Bengal, India (Teleostei: Cypriniformes: Cyprinidae). Zootaxa, 2277: 53-60.

68. Britz, R., Conway, K. W. & Rüber, L. (2009). Spectacular morphological novelty in a miniature cyprinid fish, Danionella dracula n. sp. Proceedings of the Royal Society, B, 276: 2179-2186; published on-line 11 March 2009 (doi: 10.1098/rspb.2009.0141)

67. Britz, R. & Conway, K. (2009). Osteology of Paedocypris, a miniature and highly developmentally truncated fish (Teleostei: Ostariophysi: Cyprinidae). Journal of Morphology, 270: 389-412. Published on-line 23 Dec 2008 (DOI: 10.1002/jmor.10698)

66. Britz, R. (2008). Review of FRESHWATER FISHES OF MEXICO by R. R. Miller, University of Chicago Press, USA, 2005. Acta Zoologica 89: 87-88.

65. Britz, R. (2008). Out of water (Review of YOUR INNER FISH by Neil Shubin, Penguin Books, UK, 2008). Times Literary Supplement (TLS) September 12 2008: 12.

64. Kullander S.O. & Britz, R. (2008). Puntius padamya, a new species of cyprinid fish from Myanmar (Teleostei: Cyprinidae). Electronic Journal of Ichthyology, 2008 (2): 56-66.

63. Britz, R., Kottelat, M. (2008). Paedocypris carbunculus, a new species of miniature fish from Borneo (Teleostei: Cypriniformes: Cyprinidae). Raffles Bulletin of Zoology 56: 415-422.

62. Britz, R. (2007, published 2008). Polycentridae in Poissons d’eaux douces et saumâtres de basse Guinée, ouest de l’Afrique centrale, The Fresh and Brackish Water Fishes of Lower Guinea, West-Central Africa (eds. Teugels, G. G., Stiassny, M. L. J. & Hopkins, C.), Vol. 2. Published by IRD, MRAC, AMNH, pp. 248-250.

61. Britz, R. (2007). Polypteridae in Poissons d’eaux douces et saumâtres de basse Guinée, ouest de l’Afrique centrale, The Fresh and Brackish Water Fishes of Lower Guinea, West-Central Africa (eds. Teugels, G. G., Stiassny, M. L. J. & Hopkins, C.) Vol. 1. Published by IRD, MRAC, AMNH, pp. 168-173.

60. Britz, R. (2007). Channa ornatipinnis and C. pulchra, two new species of dwarf snakeheads from Myanmar (Teleostei: Labyrinthici: Channidae). Ichthyological Exploration of  Freshwaters 18: 333-342.

59. Conway, K. & Britz, R. (2007). Sexual dimorphism of the Weberian apparatus and pectoral girdle in Sundadanio axelrodi (Ostariophysi: Cyprinidae), a miniature cyprinid from South East Asia. Journal of Fish Biology, 71: 1562-1570.

58. Britz, R. (2007). Two new species of Mastacembelus from Myanmar (Teleostei: Symbranchiformes: Mastacembelidae). Ichthyological Exploration of Freshwaters, 18: 257-268.

57. Rüber, L., Kottelat, M., Tan, H. H.,  Ng, P. K. L.& Britz, R. (2007). Evolution of miniaturization and the phylogenetic position of Paedocypris, comprising the world’s smallest vertebrate. BMC Evolutionary Biology 2007, 7:38 doi:10.1186/1471-2148-7-38

56. Britz, R. & Moritz, T. (2007). Reinvestigation of the osteology of the miniature African freshwater fishes Cromeria and Grasseichthys (Teleostei, Gonorynchiformes, Kneriidae), with comments on kneriid relationships. Mitteilungen aus dem Museum für Naturkunde Berlin, Zoologische Reihe, 83: 3-42.

55. Hilton, E., Britz, R., Johnson, G.D. & Forey, P. (2007). Clarification of the occipito-vertebral region of Arapaima gigas (Osteoglossomorpha: Osteoglossidae) through developmental osteology. Copeia, 2007: 218-227.

54. Britz, R. & Maclaine, J. (2007). A review of the eel-loaches, genus Pangio, from Myanmar (Teleostei: Cypriniformes: Cobitidae). Ichthyological Exploration of Freshwaters, 18: 17-30.

53. Britz, R. (2006). Review of EARLY STAGES OF ATLANTIC FISHES. Edited by W. J. Richards. Vol. I, 1335 pp., Vol. II, 1337-2640 pp. Published by Taylor and Francis (CRC Marine Biology Series), Boca Raton, Florida, USA, 2006. Journal of Fish Biology 69: 322-323.

52. Britz, R. (2006). Review of FISHES OF THE WORLD. By J. S. Nelson. 610 pp. 4th edition. Published by John Wiley & Sons Inc., 2006. Journal of Fish Biology 69, 1900-1901.

51. Britz, R. (2006). Review of  “STUDY OF THE DORSAL GILL-ARCH MUSCULATURE OF TELEOSTOME FISHES, WITH SPECIAL REFERENCE TO THE ACTINOPTERYGII”. V. G. Springer and G. D. Johnson with illustrations by Karolyn Darrow and the Appendix, Phylogenetic Analysis of 147 Families of Acanthomorph Fishes Based Primarily on Dorsal Gill-arch Muscles and Skeleton, by V. G. Springer and T. M. Orrell. Copeia, 2006: 323-329.

50. Hoffmann, M. & Britz, R. (2006). Ontogeny and homology of the neural complex of otophysan Ostariophysi. Zoological Journal of the Linnean Society, 147: 301-330.

49. Moritz, T., Britz, R. & Linsenmair, K. E. (2006). On the ecology of Cromeria occidentalis (Gonorynchiformes, Teleostei). Ichthyological Exploration of Freshwaters, 17: 173-180.

48. Rüber, L., Britz, R. & Zardoya, R. (2006). Molecular phylogenetics of labyrinth fishes, evolution of their breeding behaviour and remarks on their historical biogeography (Perciformes; Anabantoidei). Systematic Biology, 55: 374-397.

47. Britz, R. & Hoffmann, M. (2006). Ontogeny and homology of the claustra in otophysan Ostariophysi (Teleostei). Journal of Morphology, 267: 909-923. Published on-line April 2006, DOI: 10.1002/jmor.10447

46. Moritz, T., Britz, R. & Linsenmair, K. E. (2006). Cromeria nilotica and C. occidentalis, two valid species of the African freshwater fish family Kneriidae (Gonorynchiformes, Teleostei). Ichthyological Exploration of Freshwaters, 17: 65-72.

45. Kottelat, M., Britz, R., Tan, H. H.& Witte K.-E. (2006). Paedocypris, a new genus of cyprinid fish from threatened Southeast Asian peat swamps comprising the world's smallest vertebrate. Proceedings of the Royal Society London B, 273: 895-899. doi:10.1098/rspb.2005.3419

44. de Carvalho, M.R.,  Bockmann, F.A., Amorim, D. S.,  de Vivo, M. de Toledo-Piza, M., Menezes, N.A, de Figueiredo, J.L., Castro, R.M.C., Gill, A. C., McEachran, J. D. Compagno, L. J. V., Schelly, R. C., Britz, R., Lundberg, J. G., Vari, R. P. & Nelson, G. (2005). Revisiting the Taxonomic Impediment. Science, 307: 353.

43. Ferraris C. & Britz, R. (2005). A new diminutive species of Glyptothorax (Siluriformes: Sisoridae) from the upper Irrawaddy River basin, Myanmar, with comments on sisorid and erethistid phylogenetic relationships. Ichthyological Exploration of Freshwaters, 16: 375-383.

42. Britz, R. & Johnson, G. D. (2005). Occipito-vertebral fusion in the ocean sunfishes (Teleostei: Tetraodontiformes: Molidae) and its phylogenetic implications. Journal of Morphology, 266: 74-79. Published Online: 25 Aug 2005, DOI: 10.1002/jmor.10366

41. Johnson, G.D. & Britz, R. (2005). A description of the smallest Triodon on record (Teleostei: Tetraodontiformes: Triodontidae). Ichthyological Research, 52: 176-181.

40. Moritz, T. & Britz, R. (2005). Ontogeny and homology of the basipterygoid articulation in Pantodon buchholzi (Teleostei: Osteoglossomorpha). Zoological Journal of the Linnean Society, 144: 1-13.

39. Johnson, G. D. & Britz, R. (2005). Leis's conundrum: ontogeny and homology of the clavus of the ocean sunfishes (Teleostei, Tetraodontiformes, Molidae). Part 2 Ontogeny of the median fins and axial skeleton of Ranzania laevis. Journal of Morphology, 266: 11-21. Published Online: 17 Nov 2004, DOI: 10.1002/jmor.10242

38. Britz, R. & Johnson, G. D. (2005) Leis's conundrum: ontogeny and homology of the clavus of the ocean sunfishes. Part 1 Ontogeny of the median fins and axial skeleton of Monotrete leiurus (Teleostei, Tetraodontiformes, Tetraodontidae). Journal of Morphology, 266: 1-10. Published Online: 17 Nov 2004, DOI: 10.1002/jmor.10243

37. Britz, R. (2004). Teleostei in Westheide, W. & Rieger, R.: Spezielle Zoologie, Teil 2: Wirbeltiere. Gustav Fischer Verlag, pp. 238-285. (Chapter Teleostei in Volume 2 [Vertebrate systematics] of the major German zoology textbook).

36. Rüber, L. , Britz, R., Kullander, S. O., Zardoya, R. (2004). Evolutionary and biogeographic patterns of the Badidae (Teleostei: Perciformes) inferred from mitochondrial and nuclear DNA sequence data. Molecular Phylogenetics and Evolution, 32: 1010–1022

35. Britz, R. (2004). Egg structure and larval development of Pantodon buchholzi Peters 1877 (Teleostei: Osteoglossomorpha), with a review of data on reproduction and early life history in other osteoglossomorphs. Ichthyological Exploration of Freshwaters, 15: 209-224.

34. Kottelat, M., Kullander, S.O., Fang Fang, Britz, R. & Ferraris C. (2004). Comment on the proposed conservation of the specific name of Macropodus concolor Ahl, 1937 (Osteichthyes, OSPHRONEMIDAE). Bulletin of Zoological Nomenclature, 61: 114-116.

33. Britz, R. (2004). Polypterus teugelsi, a new species of bichir from the Upper Cross River system in Cameroon (Actinopterygii: Cladistia: Polypteridae). Ichthyological Exploration of Freshwaters, 15: 179-186.

32. Rüber, L., Britz, R., Tan, H.H., Ng, P. K. L., Zardoya, R. (2004). Evolution of mouthbrooding and life-history correlates in the fighting fish genus Betta. Evolution, 58: 799-813.

31. Courtenay, W. R., Williams, J. D., Britz, R., Yamamoto, M. N. & Loiselle, P.V. (2004). Identity of introduced snakeheads (Pisces, Channidae) in Hawai’i and Madagascar, with comments on ecological concerns. Occasional Pappers of the Bishop Museum, 77: 1-13.

30. Britz, R. (2003). Suborder Anabantoidei: Labyrinth fishes in Hutchins, M.: Grzimek's Animal Life Encyclopedia Vol. 5, Fishes II. Gale Group, pp. 427-436.

29. Britz, R. & Bartsch, P. (2003). The myth of dorsal ribs in gnathostome vertebrates. Proceedings of the Royal Society London B (Supplement), Biology Letters, 270: S1-S4. Published online 30 May 2003 (DOI: 10.1098/rsbl.2003.0035).

28. Britz, R. & Kullander, S. O. (2003). Polycentridae- Leaffishes, pp. 603-604 in Reis, R. E., Kullander, S. O., Ferraris, C. J. Jr.: Check-list of the freshwater fishes of South and Central America, EDIPUCRS: Porto Alegre.

27. Britz, R. & Johnson, G. D. (2003). On the homology of the posteriormost gill arch in polypterids (Cladistia, Actinopterygii). Zoological Journal of the Linnean Society, 138:  495-504.

26. Britz, R., Favorito-Amorim, S. & Johnson, G. D. (2003). The hyopalatine arch of a 25 mm larva of Synbranchus and homology of the pterygoid bone in the Synbranchidae (Teleostei: Synbranchiformes). Proceedings of the Biological Society Washington, 116: 337-340.

25. Britz, R. (2003). Danionella mirifica, a new species of miniature fish from Upper Myanmar (Ostariophysi: Cyprinidae). Ichthyological Exploration of Freshwaters, 14: 217-222.

24. Britz, R. & Ferraris, C. J. Jr. (2003). A new species of the Asian catfish genus Pseudolaguvia from Myanmar (Teleostei: Ostariophysi: Siluriformes: Erethistidae). Zootaxa, 388: 1-8.

23. Britz, R. (2002). Larvale Anheftungsorgane bei Teleostei in: Greven, H. & Riehl, R. (eds.): Verhalten der Aquarienfische (2). Birgit Schmettkamp Verlag, Bornheim, pp. 17-35. (Larval attachment organs in teleosts)

22. Britz, R. & Johnson, G. D. (2002). "Paradox Lost": Skeletal ontogeny of Indostomus paradoxus and its significance for the phylogenetic relationships of Indostomidae (Teleostei, Gasterosteiformes). American Museum Novitates, 3383, 1-43.

21. Britz, R. & Kottelat, M. (2002). Parasphaerichthys lineatus, a new species of labyrinth fish from southern Myanmar (Teleostei: Osphronemidae). Ichthyological Exploration of Freshwaters, 13: 243-250.

20. Kullander, S. O. & Britz, R. (2002). Revision of the family Badidae (Teleostei: Perciformes), with description of a new genus and ten new species. Ichthyological Exploration of Freshwaters, 13: 295-372.

19. Britz, R. & Cambray, J. (2001). Structure of egg surfaces and attachment organs in anabantoids. Ichthyological Exploration of Freshwaters, 12: 267-288.

18. Britz, R. (2001). The genus Betta - monophyly and intrarelationships, with remarks on the subfamilies Macropodinae and Luciocephalinae. Ichthyological Exploration of Freshwaters, 12: 305-318.

17. Kerle, R., Britz, R. & Ng, P. K. L. (2000) Habitat preference, reproduction and diet of the earthworm eel, Chendol keelini (Teleostei: Chaudhuriidae). Environmental Biology of Fishes, 57: 413-422.

16. Britz, R., Heydt, A. & Kirschbaum, F. (2000). Observations on the structure of larval attachment organs in three species of gymnotiforms (Teleostei: Ostariophysi). Acta Zoologica, 81: 57-68.

15. Breining, T. & Britz, R. (2000). Egg surface structure of three clingfish species, using scanning electron microscopy (Teleostei: Gobiesocidae). Journal of Fish Biology, 56: 1129-1137.

14. Kottelat, M. & Britz, R. (2000). Chaudhuria fusipinnis, n. sp. in: Kottelat, M.: Diagnoses of a new genus and 64 new species of fishes from Laos (Teleostei: Cyprinidae, Balitoridae, Bagridae, Syngnathidae, Chaudhuriidae and Tetraodontidae). Journal of South Asian Natural History, 5: 78-79.

13. Britz, R. (2000) Aspects of the reproduction and development of Indostomus paradoxus Prashad & Mukerji 1929 (Teleostei, Gasterosteiformes). Ichthyological Exploration of Freshwaters, 11: 305-314.

12. Kullander, S. O., Britz, R. & Fang, F. (2000). Pillaia kachinica, a new chaudhuriid fish species from Myanmar, with observations on the genus Garo (Teleostei: Mastacembeloidei: Chaudhuriidae). Ichthyological Exploration of Freshwaters, 11: 327-334.

11. Britz, R. & Kottelat, M. (1999). Carinotetraodon imitator, a new freshwater pufferfish from India (Teleostei: Tetraodontiformes). Journal of South Asian Natural History, 4: 39-47.

10. Ng, H. H., Ng, P. K. L. & Britz, R. (1999). Channa harcourtbutleri (Annandale, 1918): a valid species of snakehead (Perciformes; Channidae) from Myanmar. Journal of South Asian Natural History, 4: 57-64.

9. Britz, R. & Kottelat, M. (1999). Sundasalanx mekongensis, a new species of clupeiform fish from the Mekong basin (Teleostei: Sundasalangidae). Ichthyological Exploration of Freshwaters, 10: 337-344.

8. Britz, R. & Kottelat, M. (1999). Two new species of gasterosteiform fishes of the genus Indostomus (Teleostei: Indostomidae). Ichthyological Exploration of Freshwaters, 10: 327-336.

7. Gemballa, S. & Britz, R. (1998). Homology of intermuscular bones in acanthomorph fishes. American Museum Novitates, 3241: 1-25.

6. Britz, R. & Bartsch, P. (1998). On the reproduction and early development of Erpetoichthys calabaricus, Polypterus senegalus, and P. ornatipinnis (Actinopterygii: Polypteridae). Ichthyological Exploration of Freshwaters, 9: 325-334.

5. Bartsch, P. & Britz, R. (1997). A single micropyle in the eggs of the most basal actinopterygian fish, Polypterus (Actinopterygii, Polypteriformes). Journal of Zoology, 241: 589-592.

4. Britz, R. (1997). Egg surface structure and larval cement glands in nandid and badid fishes (Teleostei, Percomorpha), with remarks on phylogeny and zoogeography. American Museum Novitates, 3195: 1-17.

3. Britz, R. (1996). Ontogeny of the ethmoidal region and hyopalatine arch in Macrognathus pancalus (Teleostei, Mastacembeloidei), with critical remarks on mastacembeloid inter- and intrarelationships. American Museum Novitates, 3181: 1-18.

2. Britz, R., Kokoscha, M., Riehl, R. (1995). The anabantoid genera Ctenops, Luciocephalus, Parasphaerichthys and Sphaerichthys as a monophyletic group: evidence from egg surface structure and reproductive behaviour. Japanese Journal of Ichthyology, 42: 71-79.

1. Britz, R. (1994). Ontogenetic features of Luciocephalus (Perciformes, Anabantoidei), with a revised hypothesis of anabantoid intrarelationships. Zoological Journal of the Linnean Society 112: 491-508.